Sanemi and Iguro head to a mission at an abandoned castle on top of a mountain. Meanwhile, Tanjiro is recovering from the injuries he sustained in the Swordsmith Village at the Butterfly Mansion. During this time, the Hashiras gather at the Ubuyashiki residence for a meeting. Muichiro reveals the method of manifesting the marks. To enhance the overall strength of the Demon Slayer Corps, the Hashira Training begins. 实弥与伊黑等人出发到山上的废城进行任务。另一方面，炭治郎在蝴蝶

Kagaya's crow suddenly visits Tamayo. He is inviting Tamayo, a demon, to the Demon Slayer Corps' headquarters. Meanwhile, at the Butterfly Mansion, Tanjiro receives a letter from Kagaya and visits Giyu, who has not been participating in training. Tanjiro persistently talks to Giyu even though he is rejected at first. Eventually, Giyu gives in and starts to explain why he refused to participate in the training. 耀哉的鎹鸦突然出现在珠世面前，邀请身为鬼的她到鬼杀队的据点。另一方面，炭

Having recovered from his injuries, Tanjiro visits Uzui, the former Hashira, to take the Hashira Training. After a brief reunion, he quickly begins his training to improve his basic fitness. 伤势恢复的炭治郎为了参加柱训练，首先去拜访前音柱宇髓的住处。短暂地感受了久别重逢的喜悦，马上开始接受增加基础体力的训练。

After overcoming Uzui's training, Tanjiro visits the Tokito residence next. The arduous nature of Muichiro's training and his harsh words had already demoralized the Corps members who had started the training. Hearing their voices, Tanjiro is wondering if something could be done. 炭治郎结束宇髓的训练之后，接着拜访时透家。由于无一郎的训练很辛苦，讲话又严厉，已经开始训练的鬼杀队员们纷纷感到意志消沉。听到队员们询问能不能想想办法之后，炭治郎决定――。

After completing Mitsuri's training, Tanjiro moves on to Iguro's training. Upon arriving, he is met by an angry Iguro. Will Tanjiro be able to endure the terrifying training aimed at improving his swordsmanship? 结束蜜璃训练的炭治郎接着前往参加伊黑的训练。伊黑在训练场等着炭治郎，当他一抵达就大发雷霆。究竟炭治郎能否完成为了矫正刀法进行的恐怖训练呢――。

Tanjiro and Zenitsu head towards the Himejima's training grounds. There, they reunite with Inosuke and Murata. Tanjiro begins his training under the waterfall, struggling with the coldness and weight of the water. 炭治郎与善逸前往位于深山中的悲鸣屿训练场，他们在那边与伊之助跟村田再会。炭治郎首先开始冲打瀑布的训练，然而水的冰冷以及冲劲让他陷入苦战。

During the Stone Hashira's training, Tanjiro receives advice from Genya and manages to move the rock, but he collapses from exertion. Himejima steps in to help him. Touched by Tanjiro's sincerity, Himejima shares his own past with him. 岩的训练中，炭治郎听了玄弥的建议之后成功推动岩石，然而他用尽全力累倒在地，此时悲鸣屿前来援助。悲鸣屿被炭治郎直率的心意感动，开始说起自己的过去――。

On a quiet night with the moon shining brightly, Muzan Kibutsuji finally encounters Kagaya Ubuyashiki at the Ubuyashiki residence. Driven by a single desire to defeat Muzan, Kagaya clings to life, while Muzan dreams of immortality. Bound by a thousand years of fate, what will these two have to say to each other? 在一个月亮高挂的宁静夜晚，耀哉终于遇见了出现于产屋敷府的鬼舞辻?无惨。耀哉一心只想着要打倒无惨而将生命连系起来，无惨则是做着不灭的梦。有着持续千年漫长因缘的两人，将会诉说怎样的故事呢――。